Here is a roundup of major Syrian historical sites or monuments that have been destroyed by all sides in the country`s crisis.
According to the Association for the Protection of Syrian Archeology (APSA), more than 900 monuments and archeological sites have been looted, damaged or destroyed.
Referring to satellite images, the United Nations said in December that 300 sites had been plundered, damaged or destroyed.The UN training and research agency UNITAR has said satellite images "confirm the destruction of the main Temple of Bel as well as a row of columns in its immediate vicinity."
That was the second temple in Palmyra to be destroyed by the Islamic State group in a week, following one in Baalshamin.
In July, IS destroyed the statue of the Lion of Athena, which stood more than three metres (10 feet) high in front of the Palmyra museum. IS transformed the museum into a court and jail.
On August 18, IS murdered the 82-year-old retired head of antiquities in Palmyra, Khaled al-Assad, hung his mutilated body in public.
IS condemns pre-Islamic religious works as idolatry and has targeted Palmyra, which is northeast of Damascus, for destruction. The city has been listed as a world heritage site by UNESCO.
The city sustained damage in 2013 during fighting between rebels and government forces, and troops did additional harm there last year.
The historic centre of Aleppo, northern Syria, is divided between rebels and loyalists. In September 2012, a blaze swept through ancient shops in the city`s souk, or market place, and in April 2013, the minaret of the historic Omayyades mosque collapsed during fierce fighting.
Rebels using explosives to reach government positions in the old city destroyed the iconic Carlton Hotel on May 8, 2014.
In July 2015, a blast destroyed part of the ramparts that surround the citadel, a leading example of medieval Islamic military architecture. The Syrian Observatory for Human Rights said rebels had dug a tunnel under the wall, and that government troops blew it up to protect their positions.
A heavily fortified Crusader citadel known as the Crax sed Chevaliers and used as a rebel base near Homs, central Syria, has suffered damage from army shelling.On June 20, 2015, the best-known mosaic museum in Syria was seriously damaged by barrel bombs launched by regime aircraft, APSA said.In this northern IS bastion, much of the Sufi Muslim Uwais al-Qarani Mosque and a shrine to Ammar bin Yasir, a companion of the Muslim Prophet Mohammed, was destroyed, with non-governmental organisations pointing the finger at the IS.
In the northeastern region of Hassake, IS has also destroyed Assyrian statues dating from the first century, APSA said.
Other notable sites damaged or looted include Dura-Europos, Apamea, Ebla, Mari, Ajaja and Hamam Turkoman.