Brussels: The European Commission called on member states to abide by common border rules on Thursday, a day after Denmark closed a highway and rail links with Germany to stem the flow of refugees heading north to Sweden.

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The Commission was in contact with Danish authorities and following the situation closely, a Commission spokeswoman told a news conference.

"Of course we do understand that in the meantime the situation has been normalised," she said.

"This is a refugee crisis that affects all of the member states and it is not now the time for individual or isolated actions. Now is the time for a coordinated European response. And of course we can only have a well-functioning Union if all member states play by the rules." 

The motorway connecting the Danish peninsular Jutland to Germany reopened on Thursday, although some limits on rail traffic remained.